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Amphibia: Anura - Leiuperidae, Puddle Frogs
The species in the Leiuperidae were long associated with the family Leptodactylidae. Molecular evidence suggests that the 7 genera and 75 species belong in their own family. Puddle Frogs range from southern Mexico southward through Central and South America to Chile and Argentina. Grant, et al. (2006, Bulletin of the American Museum of Natural History, 299) have removed these frogs from the Leptodactylidae and consider them to be the sister taxon to Bufonidae + Hylodidae + Dendrobatoidea.
puddle frog, Engystomops pustulosus

The puddle frog, Engystomops pustulosus (Cope, 1864) ranges from eastern and southern Mexico (Veracruz and Oaxaca) southward through Central America to Colombia and Venezuela. The photographs on this page were all taken in Trinidad. This species was long considered part of the genus Physalaemus.

puddle frog, Engystomops pustulosus

The puddle frog, Engystomops pustulosus (Cope, 1864). A calling male in shallow water.

puddle frog, Engystomops pustulosus

The puddle frog, Engystomops pustulosus (Cope, 1864). A calling male in shallow water.

puddle frog, Engystomops pustulosus

The puddle frog, Engystomops pustulosus (Cope, 1864). A pair in amplexus. Amplexing pairs will on occasion leave the water and wander overland, like these individuals. They are also easy prey for the large terrestrial crabs that share their habitat.

foam nest, puddle frog, Engystomops pustulosus

The puddle frog, Engystomops pustulosus (Cope, 1864). A foam nest is made from proteins released during amplexus. The foam nest is made by the kicking legs of the frogs and the nest houses and protects the eggs from predation and desiccation.

   
   
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